Dioecious perennial herb up to 20(–30) m long, trailing or climbing with simple tendrils, with tuberous rootstock; stem green and longitudinally ribbed when young, becoming white-spotted when older and eventually woody. Leaves alternate, simple; stipules absent; petiole 1–5 cm long; blade broadly ovate to pentagonal or orbicular in outline, 3–12 cm × 3–15 cm, shallowly to deeply palmately 3–5-lobed, cordate at base, margin entire or sinuate and often with distinct reddish glandular teeth, glabrous, punctate. Flowers axillary, unisexual, 5-merous, with tubular receptacle 3–7 mm long, sepals linear, up to 6 mm long, corolla campanulate, with lobes up to 2 cm × 1.5 cm, yellow-orange; male flowers solitary or paired, rarely 3–4 in a short raceme, pedicel 1–7 cm long, stamens 3, united into a column; female flowers solitary, pedicel up to 2.5 cm long, ovary inferior, cylindrical, up to 1.5 cm long, 1-celled, style 3 mm long, stigma 3-lobed. Fruit an ellipsoid or rarely spherical berry 3–7 cm × 1–3.5 cm, fleshy, green with white stripes when young, turning red at maturity, many-seeded. Seeds asymmetrically pear-shaped in outline, compressed, c. 6 mm × 3 mm, margin rather thick and grooved. |