Large, dioecious tree up to 50 m tall; bole straight, cylindrical, branchless for up to 25(–30) m, up to 2.5(–3) m in diameter, buttresses absent or small; surface roots often long and prominent, red-brown with yellow lenticels; outer bark grey to dark brown or black, lenticelled, becoming scaly, inner bark thick, fibrous, cream-coloured speckled with orange-brown spots, exuding a white or yellowish latex; crown spreading; branches obliquely ascending. Leaves distichously alternate, simple; stipules free, slightly clasping the stem, up to 5 cm long, caducous; petiole 1–6 cm long; blade oblong to elliptical, 6–20(–33) cm × 3.5–10(–15) cm, base cordate to obtuse, often very unequal, apex acuminate, margin almost entire to wavy but toothed in juvenile specimens, papery to leathery, above glabrous or slightly hairy on the main veins, below densely short-hairy between the finest veins, pinnately veined with 10–22 pairs of lateral veins. Inflorescence a catkin, usually solitary in leaf axils or on leafless nodes at the base of twigs, white hairy, flowers numerous in longitudinal rows alternating with rows of bracts; male inflorescence 8–20(–32) cm × 0.5–1 cm, hanging, peduncle 0.5–2.5 cm long; female inflorescence 2–4 cm × 0.5–2 cm, peduncle 0.5–2 cm long. Flowers unisexual, 4-merous, sessile; male flowers c. 1.5 mm long, white, tepals 4, basally fused, stamens 4 and inflexed in bud, rudimentary pistil present; female flowers 2–3 mm long, with 4 basally fused tepals, greenish, ovary superior, c. 1 mm long, 1-celled, stigmas 2, one 3–7 mm long, the other up to 1 mm long. Fruit an ellipsoid, compressed achene 2.5–3 mm long, arranged in infructescences up to 5 cm × 1.5 cm. Seed c. 2 mm long. |